the Guangxi Natural Science Foundation(2025GXNSFHA069094);the Baise City Science and Technology Plan Project (Science and Technology Infrastructure Support Program)(KFKT02-004);the Guangdong Provincial Key Laboratory of Research and Development in Traditional Chinese Medicine(2025ZJ0707);the High-level Talent Introduction Project of Youjiang Medical College of Nationalities(yy2021sk002);the Open Project of Guangxi Key Laboratory of Medical Genetics and Genomics Research(GXKMGG202201);the Innovation Project of Youjiang Medical University For Nationalities Graduate Education(YXCXJH2023016);the Guangxi Zhuang Autonomous Region Regional Student Innovation and Entrepreneurship Training Programme Project(S202310599056)